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They're soul mates who've never met, a man and woman with parallel lives for whom first love is long gone and perfect lovers are yet to appear. Gil is an American comedy writer who came to the English countryside for what he believed was true love, while Alice is a perfume account manager for a London department store, set to move from the country and her pricey Brighton flat because of the commute. Milly and Cleo are forever trying to sniff out Mr. right for Alice, but can they orchestrate a meeting?

Alice's birthday brings presents and dates she didn't ask for, and Gil somehow gets more involved in Irene's life than he'd like.

Alice's part in a company presentation offends her, but can she stand up to a superior whose review could determine her future at the store? Meanwhile, Gil tries a blind date with a fan, but is distracted by concerns for Irene and her date.