In an attempt to stop evil Coop from destroying their dimension, Coop and Gorrath, assisted by alternate Jamie, launch a desperate assault on the evil Coop.

In the last days of the Earth War, Jamie and Kiva were busy shutting down the Glorft Core Destroyer, while Coop and the Glorft, led by Gorrath as usual, were fighting out. However, Coop and Gorrath get transported to a mirror dimension - because of a gamepad combo that activated the Transdimension Device - where Coop meets his alternate self, a futuristic, evil, and muscular warlord version of himself at that, having defeated the Glorft, abandoned Megas, formed an Empire, and keeps on destroying - no matter the location, time, dimension and enemy. He also sees that Kiva has turned cyborg as well and is evil Coop's sidekick. Alternate Jamie, on the other hand, has lost his cowardice forever and became strong enough to try and defeat evil Coop and his empire with his Resistance (and on a side note, Jamie and Kiva were dating in the future). Coop and Gorrath must work together this time to stop mirror Coop and prevent him from ever destroying Earth again.

Coop accidentally teleports himself directly into the Karrajor, the Glorft Mothership, and now Kiva and Jamie have to figure out how to pilot Megas to save him. They do okay at first, but eventually crash into the Glorft mothership and get themselves - and Megas - captured. Fortunately, Gorrath foolishly lets Coop get the car, so Coop tears through the ship, intent on rescuing his friends and Megas. Now Coop must rescue everyone while finishing off the Glorft invasion once and for all, but how is he supposed to do that when the sheer amount of Glorft mech's overpowers even Megas and tears off several vital parts? Even worse, the Glorft have a new mech, the Karrajor in Mech mode, which is even larger than the UMD, which raises the question: how will Coop defeat this new threat with a critically damaged Megas?

After racking up multiple driving violations, Coop has to spend the day at the DMV to get Megas out of impound.

The S-Force erroneously attack Coop because they believe he is an evil-doer intent on destroying the Earth; to complicate matters, Coop is aided by the S-Force's arch-nemesis, Ender, as he too believes Coop is evil.

Coop finds an alien robot in a Jersey City junkyard, and decides to soup it up with hot-rod parts and flame decals. What happens when the original owner of this robot sees Coop's improvements?

In order to impress a girl, Jamie borrows Coop's car to take her for a ride. Unfortunately, Warmaster Gorrath arrives piloting a deadly replica of Megas (dubbed Mecha-Megas) who's destroying Jersey searching for Megas; naturally, everyone thinks that the fake is just Coop being destructive. So Coop must use his secondary command bridge to control Megas. However, this method is much more difficult, so Coop does rather badly. Even worse, Jamie accidentally activates the car's connection to Megas, so now he's in control of Megas, but he has no idea - he's just trying to have a date.

Coop begins a bad day by accidentally creating a giant cheese monster and has to stop it, but the method he uses gets cheese in Megas's CPU core. Unaware of this, Coop enters Megas in an auto show, and is putting up a very good showing until Megas breaks down and nearly destroys the building. Fortunately, he's able to stop it, but now the Glorft see an opening to attack and invade the city, hoping to capture Megas. But first they need to find the CPU core.

Once again, Coop mistakenly releases a giant, plant-type alien menace that wants to destroy Earth and acquire its heat. Fortunately, Megas is too big for the plant to handle, so it has to find a bigger form. Unfortunately, this is Thanksgiving, and the spore fuses with a bunch of parade balloons to this end. Coop now must prevent the plant from wrecking the holiday.

Coop, Jamie and Kiva teleport themselves to a planet with brainwashed robots who act like sissies and workers. The alien who did it tries to do the same to Megas, but his brainwasher device only works on sentient robots. (Obviously, Coop's doesn't count.) Coop tries to put the alien out of business for good without unnecessarily destroying the prisoners. Coop in the end destroys the mind control device, freeing the robots. He then leaves the planet, unfortunatly as the group leaves it's shown that the planet was actually a prison to the worst robots in the universe. Who start destroying the planet after they leave.

The S-Force has been captured by an evil villain named Zarek, and he is planning to execute them. Fortunately, Jax escapes and manages to enlist Coop's help. Coop manages to save the S-Force from the "infinity zone," which Coop, Jamie, Kiva, the S-Force, and Megas fall into, but survive the Infinity Zone after Coop helps everyone escape, but now Zarek is sending down an entire army to terminate them. It's Megas and the Ultra-Dimensional Power Zorp vs. a bunch of insect robots! The "gift" that Zarek offers Coop, Jamie and Kiva is an end to their lives, whereas the "gifts" Coop tries to give Zarek, but almost destroys the S-Force with, is actually thousands of missiles being fired at Zarek, due to the new Super Destructor Mode.

Coop crashlands on an icy planet and ends up destroying its only mechanized guardian. Now he must protect the Yetis there from selfish, ice-hungry robots named the Cerilians, who use it to maintain their cooling systems. Trouble is, Megas is iced over, and does some of the Cerilian Army's work for him. This is going to be one icy battle for the team.

Coop, Jamie and Kiva head off to Las Vegas, and inadvertently discover Area 50. Coop accidentally activates a destructive energy-leech robot, who, due to severe and fatal programming errors, sees everything as "the enemy." Coop further screws up by directing the robot to Las Vegas. Now Coop must find a way to shut down the robot permanently before Las Vegas loses all its power. Coop in the end destroying the cities power source causing the robot to deactivate. And he then buried the robot in the Grand Canyon which he had to fill with dirt.

Kiva gets some unwanted attention from Captain Warlock, a space pirate who goes for red heads, and now must make a choice between a possible time drive or two complete idiots. As usual, one wrong choice means certain death. Now Kiva must find a way to free Coop and Jamie as well as fend of the advances of one love sick space pirate and his mechanized crew.

Coop inadvertently frees the Glorft from no-space, allowing Gorrath to start his revenge. The only thing Coop is worried about, however, is babysitting cousin Skippy, who is easily bored and hates everything about Coop's house. In an effort to get him to shut up, Coop takes Skippy to the moon in Megas. (Incredibly, he is still bored.) There he discovers the Glorft is plotting to smash the moon into the Earth. Coop has to fight off the Glorft while making sure Skippy doesn't end up Glorft fodder.

Coop's builds the world's most powerful universal remote - and Skalgar (who Coop and Jamie called "School Girl"), an ignorant, alien criminal with an inferiority complex, wants it, believing its (non-existent) destructive power can boost his reputation. Coop underestimates his short opponent, and Skalgar teleports the remote to his own mech. Coop then typically destroys half of Jersey City attempting to recover the remote.