Yoshika's plans to become a doctor are put on hold after a mysterious letter from her father leads to a reunion with Sakamoto. Elsewhere, her former Strike Witch comrades take to the air to battle a new breed of Neuroi!

A menacing new breed of Neuroi looms high above the clouds, and the Strike Witches must equip themselves with experimental rocket boosters in order to reach the unprecedented altitude required to destroy their enemy!

Major Sakamoto risks her life to save the world, but the Neuroi hive proves nearly impossible to destroy. The witches are all out of magic, but Yoshika refuses to give up the fight!

Yoshika is permanently grounded after suddenly losing the ability to fly, but a prototype Striker Unit inspired by her father's letter may hold the key to putting her back in action!

As the Strike Witches prepare for their most dangerous mission yet, Major Sakamoto - the bravest warrior among their ranks - can no longer hide the fact that her magic is rapidly fading!

Yoshika makes her decision to join the Strike Witches. Lynette Bishop, another rookie witch who is 15 years old, shows Yoshika around. Although Yoshika is nervous about her new assignment and understands nothing about the arrangement of the troop, Lynette acts in a somewhat unfriendly manner, adding to Yoshika's anxiety

Yoshika gradually orients herself to life as a member of Strike Witches. However, she is condemned by her supervisor Gertrud Barkhorn as being inexperienced. Desperate to win her acceptance, Yoshika gets carried away in her training when she suddenly comes face to face with Neuroi.

On their journey to Britannia, Yoshika and the others are assaulted by mysterious invaders called the Neuroi, menacing beings from another world. In order to buy time until reinforcements arrive, Major Sakamoto sallies forth alone, only to be met by a fierce attack from the opposition. Despite her best efforts she is driven back and ultimately cornered. Yoshika can do nothing but wait in terror inside the battleship.

Barkhorn runs into big trouble during a test flight of the new jet striker unit, and only an extremely well-endowed comrade has the power to save her from the ultimate crash-landing!

Erica misplaces a key part of her uniform (panties), causing a chaotic incident in Witches Base.

Yoshika and her comrades are attacked by the Neurois on their way back from a mission in London, but gets help from Sanya who happened to be on a night patrol. As a result, Yoshika is given orders to serve the night shift, and must spend the night in the same room with Sanya, when an envious Eila barges in.

Rocket attacks from the Neuroi nest in Berlin are threatening the base in Kiel. Only Mina has what it takes to stop them, but her magic power is fading…

The girls take a detour to Nederland, to help a legendary flower bloom.

With Miyafuji's magic power disabled, humanity is forced to use its backup plan: the powerful Ratte tank. But can they win without their ultimate ace?

Yoshika is trapped inside the dome around Berlin, and the rest of the Witches are cut off. Can they get inside to save her in time?

Shirley's world record was just broken, and she's not happy about it. As Shirley and Lucchini attempt to beat the new world record, an unusual Neuroi threat swiftly approaches.

A mysterious idol causes all the witches' breasts to grow bigger and bigger, and the girls gradually fall victim one by one...

When Erica is shot down in enemy territory, Gertrud will do whatever it takes to get her back.

Operation Southwind begins with a full-scale air assault on Berlin. When Yoshika is ordered to stay behind to conserve her magic, Shizuka steps up to try and fill the hole.

Yoshika leaps back into action to save her friend's father, but her striker unit almost blows up on the tarmac!

Yoshika's lost control of her magic… again! It's time for more training and more hard work if she wants to get it back!

When a Neuroi strikes at Europe's biggest supply port on the eve of humanity's counterattack, Yoshika must hold it off alone, without any weapons at all!